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The fossil, a fragment of horse leg bone, was too old for radiocarbon dating, but Willerslev estimated, based on its location in the permafrost, that it was between 560,000 and 780,000 years old.
He and an international team of colleagues then set about scouring the fragment for any trace of collagen or other material that could harbor the ancient horse's DNA. No DNA had been salvaged and sequenced from a fossil more than 130,000 years old (that was a polar bear jawbone), and theoretical estimates put the upper limit of DNA survival at about 1 million years.
The discovery offers new insights into equine evolution and raises the prospect of recovering and exploring older DNA than previously thought possible.
"There's no question this is a landmark study," says David Lambert, an evolutionary biologist at Griffith University in Australia who was not involved in the work.
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Genes that are involved in immunity, the sense of smell, and muscle development have all changed significantly since this ancient horse roamed the Yukon, Willerslev says.
Beyond its implications for equine evolution, the study pushes the envelope for ancient DNA research, Willerslev says.